3) M/N = R/S but in this scenario, because N at the time of marking was not the actual size of the population, the R/S ratio at the end of the season will be smaller than predicted (since S will be la...3) M/N = R/S but in this scenario, because N at the time of marking was not the actual size of the population, the R/S ratio at the end of the season will be smaller than predicted (since S will be larger than expected because not all of those individuals were there at the start of the season).