Studies using quantitative trait loci (QTL) mapping have associated the following loci with the matched functions; Pln-1 and Pln-4 with onset of foraging age, Pln-1 and 2 with the size of the pollen l...Studies using quantitative trait loci (QTL) mapping have associated the following loci with the matched functions; Pln-1 and Pln-4 with onset of foraging age, Pln-1 and 2 with the size of the pollen loads collected by workers, and Pln-2 and pln-3 were shown to influence the sugar concentration of the nectar collected. Handling time (h) is the amount of time it takes the predator to handle the food, beginning from the time the predator finds the prey item to the time the prey item is eaten.