Evolvability has emerged as the connection between population-level evolution and macroevolution. Although both of these processes are fundamentally the same (both occurring through mutation, drift, a...Evolvability has emerged as the connection between population-level evolution and macroevolution. Although both of these processes are fundamentally the same (both occurring through mutation, drift, and selection), they happen at such different scales that we typically use very different tools and mindsets to study each of them.