Well-managed protected areas are essential tools for securing intact ecosystems and the biodiversity they sustain. However, it will not be sufficient to rely solely on protected areas to preserve biodiversity. Harming ecosystems such as rivers and streams on unprotected lands has repeatedly been shown to decrease biodiversity also within protected areas (Colvin et al., 2011; Woodborne et al., 2012). Furthermore, many species only occur on unprotected lands (Beresford et al. 2011), and some species even fare better outside protected areas (Murgatroyd et al., 2016). Other species need to move out of protected areas to access important seasonal resources. Lastly, restoring damaged areas and maintaining intact ecosystems inside and outside of protected areas provides ecosystem services such as water and air purification. And so, while we must continue to pursue protected areas for the many benefits they do offer, we must not forget about the value of the spaces between protected areas. In this chapter, we will explore how efforts on unprotected lands can complement conservation efforts in protected areas.